01 Reproduction in Lower and Higher Plants - part 09 - Development of Embryo
01 Reproduction in Lower and Higher Plants - part 09 - Development of Embryo
Development of Embryo :
- The process of development of zygote into an embryo is called embryogenesis.
- The embryo is developed at the micropylar end of embryo sac.
- The growth of embryo triggers only after certain amount of endosperm is formed.
- After fertilization the embryonic development begins. The zygote divides to form two- celled proembryo.
- The larger cell towards the micropyle is called basal or suspensor initial cell and smaller cell towards chalaza is called terminal or embryonal initial cell.
- The suspensor cell divides transversely in one plane to produce filamentous suspensor of 6-10 cells.
- The first cell of the suspensor towards the micropylar end becomes swollen and function as a haustorium.
- The lowermost cell of suspensor is known as hypophysis.
- The suspensor helps in pushing the embryo in the endosperm.
- The embryonal initial undergoes three successive mitotic divisions to form octant.
- The planes of divisions are at right angles to each other.
- The lower tier of four cells of octant give rise to hypocotyl and radicle where as four cells of upper tier form the plumule and the one or two cotyledons.
- The hypophysis by further division gives rise to the part of radicle and root cap.
- Subsequently, the cells in the upper tier of octant divide in several planes so as to become heart shaped which then forms two lateral cotyledons and a terminal plumule.
- Further enlargement of hypocotyl and cotyledons result in a curvature of embryo and it appears horse-shoe shaped.
- The embryo development is similar in both dicots and monocots up to the octant stage. The difference appears later.
- In monocot embryo, single cotyledon occupies terminal position and plumule is lateral.
- The single shield shaped cotyledon is called as scutellum. The protective sheath of plumule is called coleoptile and that of radicle is coleorhiza.
- Finally, ovule is transformed into the seed and ovary into the fruit.
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